Lego Skylines is far more than a mere cosmetic overlay on the established Cities: Skylines formula; it is a distinct city-building experience crafted from the ground up to blend the recognizable mechanics of the genre with the unique charm and tactile nature of Lego.

Players begin by selecting a Lego minifigure avatar, establishing a tangible presence within the game world. The initial focus is not on broad zoning but on the construction of individual homes for Lego families with specific needs and desires. Fulfilling requests, such as placing a garden gnome for a family, contributes to an Appeal Level. Housing options are presented through menus, allowing players to choose by size and then by aesthetic, with each house of a given size offering identical functionality but varied appearances, from modern city pads to rustic farmsteads. The chosen house type influences the environment residents desire around their homes. Additionally, decorative elements like plaza fountains, slides, or food trucks can be placed in square panels near houses, reflecting "flavors" such as cozy, traditional, or modern, further influencing resident satisfaction.

The game encourages granular creativity through customization menus for every building, decor item, and workplace. Players can alter roof color palettes, the types of flowers in gardens, or add features like wood sheds and benches. The act of placing and removing these elements is accompanied by a satisfying digital "snap" sound, mimicking the connection of Lego bricks, with digital pieces being easily disassembled and discarded.

Workplaces are essential for citizen Happiness, tax revenue, and resource chains. Initial workplaces include farms, with the option to expand into various industries and public services. Resources are color-coded and visible on the map during workplace construction. While matching resource colors can boost productivity, players are not strictly required to do so, a recurring theme of flexibility in Lego Skylines.

In contrast to the immediate pressures of Cities: Skylines, Lego Skylines introduces concepts like resource chains and Happiness Levels gradually, avoiding overwhelming the player early on. The initial emphasis is on creative construction rather than managing the immediate needs of a growing population. Leveling up is a deliberate process, requiring players to manually initiate it after accumulating enough points, allowing for leisurely expansion and engagement with optional side quests.

As humble homesteads evolve into varied and impressive cities, players can zoom in to observe individual minifigures. Side quests often bring players face-to-face with unique Lego minifigs, each with distinct personalities and requests. These can range from building a secluded woodland house for an off-grid enthusiast to finding escaped chickens. The game's dialogue often features recognizable Lego-style humor, and the personal nature of these requests and the minifigs' names create an immersive roleplaying experience.

While players cannot currently construct buildings from scratch, relying instead on presets, the potential for a robust modding scene is anticipated. The gameplay can be described as carefully connecting pre-built sets to form a larger creation, yet it retains a personal feel. Lego Skylines simulates city life deeply, with each minifigure having a job, family, home, and leisure preferences. Players can observe citizens walking, socializing, shopping, and engaging with placed leisure facilities like skateparks.

With its gradual introduction of mechanics and a gentle learning curve, Lego Skylines appears well-suited as an entry point to the city-builder genre for younger players. It balances relaxed construction with engaging rewards and varied challenges. Sandbox Mode offers extensive customization options, allowing for a purely creative experience. However, the game is not exclusively for children; even seasoned Cities: Skylines players may need to experiment with certain mechanics. The game demonstrates that deep simulation and relaxing city-building can coexist, raising the question of why a Lego city-builder has not been a prominent fixture until now.